Finding. The African giant shrew is a continent-wide small mammal whose mitochondrial tree does not match its nuclear genome. Dianat, Mikula and colleagues show that mismatch is repeated hybridization, not a single messy ancestor. Three major lineages, four introgression events, five still-mixing geographic groups, and expansions timed to Late Pleistocene and Early Holocene climate (and maybe people).
Score 54. Mitonuclear evolution. High confidence in the genomic claim. Not an organelle-function paper.